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Submission Preparation Checklist

As part of the submission process, authors are required to check off their submission's compliance with all of the following items, and submissions may be returned to authors that do not adhere to these guidelines.
  • All co-authors have read and agree with all editorial policies and the Privacy Statement of Plant Ecology and Evolution.
  • The manuscript has not been published, in whole or in part, in another media. The manuscript is not under consideration elsewhere (submitted to a journal, series, as part of a book…).

  • All co-authors have contributed substantially to the manuscript, have approved this submitted version of the manuscript and are aware of the submission.

  • The corresponding author takes the responsibility to keep his co-authors informed on the status of the submission and on the changes required or applied to the original version of the manuscript during all stages of the submission (registration, peer-review process, proofreading process…).
  • The manuscript and the illustrations within are original, or authors have taken all the necessary steps to avoid breach of copyright, e.g. by obtaining permission to reproduce. Authors understand that they will be held personally and financially responsible for any failure to comply with this condition. This guarantees that we can publish the manuscript without infringing anyone's copyright, and labelling correctly any material that cannot be distributed under the terms of the Creative Commons Attribution License.

  • All persons/institutions/sources of support have been acknowledged, whether they were involved in the production of the results/data presented in the manuscript, or in writing/formatting the manuscript.

  • In case new material has been collected for purposes of the study, all the necessary permits have been obtained prior to collecting. The study was undertaken in compliance with all legal obligations regarding access and benefit sharing of the biological collections used in this manuscript.

  • For taxonomic papers, the type material of new taxa names has been deposited in a public herbarium or public natural history collection where it can be consulted by other researchers.

  • The manuscript files are conform to the format required in the Author Guidelines.

  • The text and figure/table captions adhere to the stylistic and bibliographic requirements outlined in the Author Guidelines. Whenever available, the Digital Object Identifier (DOI) is provided for each bibliographic reference.

  • At least two potential reviewers are provided in the Comments to Editor, with their affiliation and email address. For a definition of “potential reviewers”, please check the Cover Letter template.

  • Any potential conflict of interest is mentioned in the cover letter, especially conflict of interest with some associate editors or some members of the editorial board of the journal (see Editorial Team for full list of names). You can also list in your cover letter the “opposed reviewers”, i.e. reviewers with whom you (or co-authors) might have a conflict of interest.

Author Guidelines

The Author Guidelines are also available in pdf format here.



Plant Ecology and Evolution is an international journal devoted to ecology, phylogenetics and systematics of all ‘plant’ groups in the traditional sense (including algae, cyanobacteria, fungi, myxomycetes), also covering related fields such as comparative and developmental morphology, conservation biology, evolution, phytogeography, pollen and spores, population biology, and vegetation studies. Submissions concerning tropical and subtropical Africa are particularly welcome.


The journal is a continuation of Belgian Journal of Botany, incorporating Systematics and Geography of Plants. It is published by Meise Botanic Garden and the Royal Botanical Society of Belgium.


The journal publishes original research articles, checklists, short communications, reviews and book reviews. Manuscripts may be of any length, but must be clearly and concisely written. A regular paper reports results of a substantial, completed and original work. A checklist presents and critically discusses a list of taxa in a given frame (e.g. floristic observations in a defined geographical area, taxa from a selected group…). A short communication is a brief report of a small experiment, taxonomic proposal, or unexplained observation. A review presents a novel and critical appraisal of a specific topic.


The journal publishes contributions in English (British spelling – Oxford English Dictionary) or French. A native speaker should check manuscripts written by non-native speakers prior to submission.


When submitting to Plant Ecology and Evolution, authors agree to the journal’s policies on Copyright and Open Access, peer review process, publication ethics and publication fees.  Authors also agree to the Privacy Statement of the journal.

Authors submit their manuscript electronically by sending an email to the editorial office, The message should include the following attached files: (1) a Cover Letter; (2) a Manuscript Text file with the entire text and figures captions; (3) a pdf wherein all figures and tables are assembled, with their captions, (4) additional separate files for figures and tables; and eventually (5) supplementary files. Please check the instructions below to prepare your files prior to submissions. All file types can be submitted as supplementary files (PDF, nexus, Excel, video files…). They will be reviewed along with the core paper, but the author is responsible for the layout of supplementary files. Authors of phylogenetic papers are encouraged to submit the aligned matrices as supplementary files.

Upon submission, the corresponding author should put all co-authors in copy of the message.

Authors should use the guidelines and examples below to prepare the submission files. There are no charges to submit and/or publish in Plant Ecology and Evolution, and the co-publishers support all costs. Therefore, to limit these costs, authors are expected to have their papers well checked for content quality, format conformity and correctness in presentation of text and illustrations. A manuscript not conform to the guidelines will be returned to the corresponding author until adequate files are provided.

In case the files are too large to be sent as attachments, the corresponding author should contact the editorial office for an alternative way to access the files.

The editorial office acknowledges receipt of the submission by email within five working days.


All contributions are subject to normal reviewing processes. The reviewing model is single-blind peer review: reviewers are aware of the authors’ identities, while authors are not informed of the reviewers’ identities. The peer-review process is detailed here.

The final decision on acceptance or rejection is made by the editor in chief, upon recommendation of the communicating editor. The editorial office will communicate this final decision by email to the corresponding author. Reviewers who participated in the process will be informed separately of the final decision.


Once a manuscript is accepted, all correspondence goes through the editorial office ( Authors are required to respond to all technical remarks from the editorial office (format, style, file types…), and to fill in a publication agreement.

The copy editor prepares the paper for publication and ensures that style and form are consistent.  Depending on the number of manuscripts already ‘in press’ and on the manuscript’s length, the delay to receive proofs can vary.

Proofs must be returned to the editorial office within five working days.


Authors are invited to submit eye-catching photographs related to the submission for possible inclusion on the cover of the issue in which their paper will appear. The technical requirements are similar to those for manuscript photographs (see checklist). The image should be sent as a TIFF file of 450 dpi, size approx. 15 × 10 cm.


Authors pay no fees or page charges. Colour photographs and graphics are accepted without charge when their use enhances scientific content or clarity. The corresponding author receives a free complimentary copy of the issue in which the paper is published and a PDF (portable document format) file of the paper.

The postal address used to send the complimentary copy is the first institutional address of the corresponding author mentioned in the manuscript. If the corresponding author wishes to receive the complimentary copy at another address, he/she must inform the editorial office before proofs’ approval.


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The following documents must be provided as separate files: Cover letter; Text file; each table; each figure; (if available, each supplementary file). Authors must also provide a pdf assembling: all tables and their captions + all figures and their captions (without the manuscript's core text).

Files types and extensions

- Cover letter, Manuscript Text file, Tables files: MS Word format (extension .doc or .docx) or Rich Text Format (extension .rtf)
- Figures files: image files; extensions allowed: .svg, .ai, .pdf (saved in high-quality), .tif, .jpg, .psd, .png. Check the figure requirements below for full technical instructions.
- Assemblage of all tables, figures and their captions: .pdf
- Supplementary files: most extensions allowed (.doc, .tif, .xls, .fas, .mov, .avi…).

Naming the files

Each file is named using the first author’s initials, followed by underscore (_) and by the type of file and eventually number, for example: “AP_cover.doc”, “AP_text.doc”, “AP_table1.doc”, “AP_fig1.tif”, “AP_fig2.jpg”, “AP_supp_file1.xls”, “AP_tables_figs.pdf”



The corresponding author provides a Cover Letter in the following cases:
- submission or resubmission of a manuscript
- during the peer-review process, when sending a revised version of the manuscript.

First submission

Authors are requested to use the MS Word Cover Letter template available on the website. By completing the form, the corresponding author engages his responsibility and the responsibility of all co-authors. If the form is not complete (e.g. some boxes unchecked), the manuscript will be rejected without review. If you cannot use MS Word, please contact the editorial office for an alternative Cover Letter form.


Authors are requested to use the same Cover Letter template available on the website. In the field “Description of the manuscript”, the corresponding author indicates the manuscript reference number, as well as the name of the communicating editor of the first submission. Authors should also use this field to describe which major changes were applied to the manuscript and how the authors addressed the reviewer’s comments after the previous submission was rejected.

Revised version

Authors are requested to prepare a Cover Letter for a revised version. This letter is in MS Word format (file extension .doc or .docx) or in Rich Text Format (file extension .rtf), and contains:
- contact details of corresponding author, title and authorship of the paper
- a detailed description of all changes made in the manuscript, as well as answers to all comments and suggestions from the communicating editor and/or from the reviewers. When suggestions have not been followed, the corresponding author must provide detailed explanations about where and why changes were not carried out.

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General requirements

The text is formatted in Times New Roman, 12 pt, single-spaced throughout, with 25 mm margins. All pages and lines are numbered consecutively.

The Text File includes tables captions and figures captions (at the end of the manuscript) but excludes tables and figures – these should be submitted as separate files (see appropriate sections below).

First page

Running title: maximum 75 characters, spaces included.

The full title is informative and concise. Authorships of taxa names are omitted in the title. A higher rank (family, order…) is indicated if the title includes generic or infrageneric names of plants. Title words are not capitalized (except proper nouns and supra-generic names of taxa). Hence: “Development and multiplexing of microsatellite markers using pyrosequencing in a tetraploid plant, Vaccinium uliginosum (Ericaceae)”, and not “Development and Multiplexing of Microsatellite Markers Using Pyrosequencing in a Tetraploid Plant, Vaccinium uliginosum (Ericaceae)”

The full names of all co-authors are listed in the correct order. The full name consists of: first name, eventually initial(s) of middle name(s), and surname. When all authors are not affiliated with the same institution, each author’s name is followed by an identifying superscript number (1,2,3…) associated with the appropriate institutional address. The corresponding author is identified with a superscript asterisk (*), inserted just after the superscript number(s).

Each institutional address(es) of each author is preceded by the appropriate superscript number.

Mention “Corresponding author:” followed by the e-mail address of the corresponding author. There can be only one e-mail address, and no phone number.

Second page

Structured abstract not exceeding 300 words. Each paragraph is introduced with an appropriate heading (e.g. ‘Background and aims’, ‘Material and methods’, ‘Key results’, ‘Conclusion’). The abstract should read independently from the manuscript, i.e. include no bibliographic references and no unexplained abbreviations.

List of maximum ten keywords. Keywords ensure accurate indexing by search engines and databases, thus are important for discoverability of your paper.

For papers in French: an English translation of the full title and the abstract.

Text structure

IMRAD structure preferred (INTRODUCTION, MATERIAL AND METHODS, RESULTS, and DISCUSSION). This structure is completed by the following sections:
- if available: SUPPLEMENTARY FILES section: paragraph with short descriptive caption for each supplementary file
- ACKNOWLEDGEMENTS: use this section to acknowledge persons (or organisations) who provided: facilities, funding (include grant numbers), permits (include permit numbers), logistic support, data sharing, scientific guidance...
- REFERENCES section: list of all bibliographic references cited in the manuscript files.

Results are presented in a concise way and data are not repeated in both graphical and tabular form.

The Discussion section avoids extensive repetition of the Results.

For most manuscripts, no more than three levels of headings are used. For specific headings in taxonomic manuscripts, please check the appropriate section below.
- Main headings are in uppercase and centred (e.g. MATERIAL AND METHODS).
- Second level headings are in bold, aligned to the left. Only the first word is capitalized (e.g. Molecular analyses and not Molecular Analyses).
- Third level headings are in bold (only first word capitalized), directly followed by a n-dash (–) and by the text (e.g. Morphological characters – Corolla…).

All tables and figures are cited in the text, and numbered consecutively. First figure is "fig. 1", second figure is "fig. 2", etc.  

When starting a sentence, use ‘Figure’, ‘Figures”, ‘Table’ or ‘Tables’; otherwise, use ‘fig.’, ‘figs’, ‘table’ or ‘tables’. Some examples:
- “Figure 1 shows…”
- “The localities are shown on fig. 2…”
- “The populations (tables 1 & 2)…”
- “The petals (figs 4A, 5E) and the stipules (figs 4B–C, 5A, 6B)…”

A figure composed of several elements (A, B, C...) remains in singular form, for example
- “Bayesian analysis (fig. 2A, B)…” [...] "The localities (see fig. 3A, D, E–G)..."

Style consistency

Font styles (bold, italic, underline) are not used for emphasis, but reserved to specific usages.

Boldface is only used in second and third level headings, for introductory terms of figures and tables captions, and to stress some taxonomic acts such as new synonymy, lectotypification (see Taxonomic Treatment section).

Italic is only used for generic/infrageneric names and in specimen citations (see Taxonomic Treatment section); for Latin expressions in full (in situ), for gene abbreviations, for standard expressions and certain statistical symbols, and for journal titles (References section).

Underline is only used to indicate the major organs in a plant description (see Taxonomic Treatment section).

Numerals:  numbers up to and including ten are spelled out, except when they are measurements (6 mm); all other numbers are written in figures (24, 53…), except at the beginning of a sentence (“Thirty-two plots…”). Numerals are followed by a single space, then by the unit (24 cm).

Dates format: day, month abbreviation, year: “10 Jan. 2016”; “from 14 Jun. to 28 Jul. 2013”; “14 Jun.–28 Jul. 2013”; month, year:  “data collected in September 2014”.

Use of n-dash:
- for all ranges (measurements, elevation range, page number range…), without spaces around, e.g. “2.5–5 mm”, “150–200 m a.s.l.”, “10–14 stamens”
- for an enclosed phrase – such as this one – within a sentence (note the spaces around n-dashes)
- n-dash should not be used if you introduce the range with a preposition: do not write “from 10–15 cm”, “between 10–15 cm” but write “from 10 to 15 cm”, “between 10 and 15 cm”.

Geographical coordinates: preferred format 49°00′51.9″N, 13°34′38.4″E (degrees minutes seconds). Use “prime” character for minutes, and “double prime” character for second, no space between each element, and a comma (followed by a space) to separate latitude and longitude. Alternative format: decimal degrees.

Footnotes should be avoided.

Standard symbols are used for:
- units of measurements: SI units and SI-derived units symbols are used throughout the manuscript, as well as some non-SI units (s, min, h, d, ha…)
- chemical elements (e.g. in molecular formulas such as NaCl, O2).

Abbreviations or acronyms are used for
- molecular terminology (e.g. bp, SDS
- common statistical terms (e.g. ANOVA, PCA, t-test, R2)
- names of chemicals (ATP, MES, MOPS, HEPES
- procedures (e.g. PCR, PAGE, RFLP)
- herbaria (e.g. P, MO, LBV: see Index Herbariorum).

Material and methods section, phylogeny

When specialized equipment is mentioned in Material and methods, details of the model and manufacturer are given.

Software should be cited as true bibliographic reference, e.g. as a website reference. If the software was downloaded at an earlier time, authors should specify it in the text. For example, in the text “FigTree, version 1.4.4 (Rambaut 2018, downloaded on 1 Dec. 2018)” and in the References section “[accessed 21 Feb. 2019]” when the website url was verified later. See the example in the References section.

Databases and data sets used as sources should be cited as true bibliographic reference. For example, cite databases such as Index Herbariorum for herbarium acronyms, The Plant List, IPNI, Tropicos, AlgaeBase as sources to verify name information, GBIF as source of specimen data sets, etc.

Novel sequences for proteins or nucleotides are deposited within one of the main databases (e.g. European Nucleotide Archive, GenBank), and accession numbers are included in the paper.

Sequence matrices are preferably provided as supplementary file(s); this is mandatory when alignment information is critical to the message of the paper.

Phylogenetic trees and alignments: independently from the submission files, authors are strongly encouraged to submit their data to TreeBASE (no charges to submit data) or Dryad (data publishing charge of $120).

Scientific names of taxa

Taxonomic nomenclature follows the rules of the most recent edition of the International Code of Nomenclature for algae, fungi and plants (Shenzhen Code).

Scientific names are written out in full (Genus species) in the abstract and again in the core text at first mention of every taxon. The genus name in full is only needed for the first species in a list within the same genus (e.g. Lolium annuum, L. arenarium). Only generic and infrageneric names are italicized. Authorship (e.g. L., Benth., H.P.Linder) is added at first mention of the taxon in the core text and in the taxonomic treatment.

Citation of authorships in non-taxonomic papers:
- the author can omit authorships in the core text if they are available elsewhere, e.g. in a table or in a supplementary file
- authorships may also be omitted if the author mentions in the Material and methods section a taxonomic work (bibliographic reference) followed.

Authorship follows the standard abbreviations of author’s names available on International Plant Names Index (, without space after first name/middle name initials (e.g. H.P.Linder, not H.P. Linder).

Additional format requirements for taxonomic treatments are available below (Taxonomic Treatment section).

References in the text

All references cited in the text are provided in the References section and vice versa; this rule applies also to the literature cited in the frame of a taxonomic treatment.

Use the following format for citations in the text:
- single author: Fisher (2012), (Fisher 2012); mention of a page number: Fisher (2012: 487), (Fisher 2012: 487)
- two co-authors: Fisher & Perry (2009), (Fisher & Perry 2009)
- three or more co-authors: Fisher et al. (2012), (Fisher et al. 2012).

Multiple references are listed in chronological order and separated by semicolons. Multiple references sharing the same authorship are grouped, years separated with commas, and inserted in the list based on the oldest reference: (Cremer 1997; Fisher 2012, 2014; Blackwell et al. 2013, 2017; Adams et al. 2014, Wright & Baker 2017).

Letters a, b, c… are added to date of papers with identical authorship and year (Fisher 2011a, 2011b; Adams & Wright 2014a, 2014b; Williams et al. 2016a, 2016b, 2016c); see the References section rules for applying letters.

Reference to a work in press is only used when the work has been accepted for publication in a named journal/book/series; otherwise, the terms ‘unpubl. res.’ (unpublished results), ‘pers. obs.’ (personal observations) or ‘pers. comm.’ (personal communication) are used, and the name and affiliation (institution at least) of the person is given (e.g. S. Fisher, University of Oxford, unpubl. res.; T. Baker, Universiteit Leiden, pers. comm.)

References to grey literature (e.g. reports) are, as far as possible, avoided.

References section

References are ordered alphabetically, based on the surname of the first or sole author. If two different first authors share the same surname, the order is based on the first name initial (and subsequently middle name initial) of each author: first “Miller J.”, then “Miller P.”, then “Miller P.W.”.

References sharing the same first author surname and initial are arranged according to the following order:
1°) single-authored papers are listed first and arranged chronologically (from the oldest to the newest year of publication)
2°) two-authored papers are then listed, arranged alphabetically based on surname of second author; when papers share the same authorship in this list, they are arranged chronologically
3°) three-or-more-authored papers are then listed, arranged chronologically.
When three-or-more-authored papers share the same year of publication, they are grouped alphabetically according to surname of second author, surname of third author, etc. In such group, the first reference is identified by letter “a”, the second reference by “b”, etc.

Example of order:
Adams B. (2015)
Adams B. (2018)
Adams B., Sellowe C. (2017)
Adams B., Sellowe C. (2019)
Adams B., Wright J.H. (2014a)
Adams B., Wright J.H. (2014b)
Adams B., Wright J.H. (2016)
Adams B., Smith R., Harley W. (2016)
Blackwell P.T., Bailey F. (2015)
Blackwell P.T., Ashworth I.S., Williams L.S. (2017a)
Blackwell P.T., Bailey F., Smith R. (2017b)
Blackwell P.T., Rogers L.M., Bailey F. (2013)
Cremer (1997)
Fisher M. (2012)
Fisher M. (2014)
Fisher M., Maxwell D. (2012)
Fisher M., Maxwell D. (2016)
Fisher M., Perry K.A. (2009)
Fisher M., Cornel D., Templeton R.W. (2012)
Williams J., Johnson R.P., Wright J.H. (2016a)
Williams J., Perry K.A., Bailey F. (2016b)
Williams J., Perry K.A., Johnson R.P. (2016c)
Wright J.H., Baker (2017)

Authors are required to provide, if available, the Digital Object Identifiers (DOIs) when they list their references. The DOI (beginning with “10.”) is generally available on the main webpage of the paper, of the book or of the book’s chapter. Authors can also use the CrossRef’s free DOI lookup tool ( The DOI is introduced with the url starter

Words in article titles are not capitalized, except the first word (or word after a colon), proper nouns, institutions, geographical locations, official denominations and suprageneric names of taxa.

Generic/infrageneric names of taxa and journal titles are italicized.

Journal titles are written in full (no abbreviations). The volume number is given, and if available, the issue number.

For book and book chapters, the following information is given at the end of the citation, before the DOI: City, Publisher.

For websites, the url is followed by the date when the resource was last accessed by the authors.

References conform to one of the following styles according to the type of publication:

- paper without Digital Object Identifier (DOI):

Carlini-Garcia L.A., van den Berg C., Martins O.S. (2002) A morphometric analysis of floral characters in Miltonia spectabilis and Miltonia spectabilis var. moreliana (Maxillarieae: Oncidiinae). Lindleyana 17(3): 122–129.

 - paper with DOI:

Baker A.J.M., Dalby D.H. (1980) Morphological variation between some isolated populations of Silene maritima With. in the British Isles with particular reference to inland populations on metalliferous soils. New Phytologist 84(1): 123–138. 

Felsenstein J. (1985) Confidence limits on phylogenies: an approach using the bootstrap. Evolution 39(4): 783–791.

Dančák M., Hroneš M., Sochor M., Sochorová Z. (2018) Thismia kelabitiana (Thismiaceae), a new unique Fairy Lantern from Borneo potentially threatened by commercial logging. PLoS ONE 13(10): e0203443.

White F. (1979) The Guineo-Congolian Region and its relationship to other phytochoria. Bulletin du Jardin Botanique National de Belgique 49: 11–55.

- thesis:

Sánchez M. (2012) Conservation genetics and biogeography of the Caribbean pine (Pinus caribaea var. bahamensis) in the Bahamas archipelago. PhD thesis, Birkbeck University of London, UK.

- chapter in book:

Arcand N.N., Ranker T.A. (2008) Conservation biology. In: Ranker T.A., Haufler C.H. (eds) The biology and evolution of ferns and lycophytes: 257–283. New York, Cambridge University Press.

Barrett S.C.H., Pannell J.R. (1999) Metapopulation dynamics and mating-system evolution in plants. In: Hollingworth P.M., Bateman R.M., Gornall R.J. (eds) Molecular systematics and plant evolution: 74–100. London, Taylor & Francis.

- book:

Manly B.F.J. (1994) Multivariate statistical methods, a primer. 2nd Edition. London, Chapman and Hall.

Lestrel P.E. (1997) Fourier descriptors and their applications in biology. Cambridge, Cambridge University Press.

- book belonging to a series, a flora:

Hallé N. (1961) Sterculiaceae. In: Aubréville A. (ed.) Flore du Gabon vol. 2. Paris, Muséum national d’Histoire naturelle.

- book chapter belonging to a series, a flora:

Sales F. (2005) Vitex. In: Pope E.V., Martins E.S. (eds) Flora Zambesiaca vol. 8(7): 73–92. Kew, Royal Botanic Gardens.

- website or online document:

Thiers B. (continuously updated) Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. Available at [accessed 26 Aug. 2018].

IUCN (2017) Guidelines for using the IUCN Red List Categories and Criteria. Version 13. Prepared by the Standards and Petitions Sub-Committee. Available at [accessed 21 Feb. 2019].

- software:

Swofford D.L. (2000) PAUP* Phylogenetic analysis using parsimony (* and other methods). Version 4. Sunderland, Sinauer Associates.

Rambaut A. (2018) FigTree. Version 1.4.4. Available at [accessed 21 Feb. 2019].

- software documentation:

ter Braak C.J.F., Šmilauer P. (2002) CANOCO reference manual and user’s guide to CANOCO for Windows: software for Canonical Community Ordination. Version 4.5. Ithaka, NY, Microcomputer Power.

Figure captions

A figure composed of several smaller subfigures is treated as a single figure; each subfigure is labelled with a capital letter (A, B, C… and after Z: AA, AB, AC…). Description of each subfigure is introduced with the capital letter (in bold) followed by a full-stop (not bold). Boldface is not used for subsequent information (e.g. repetition of capital letter for scale bar information).

In figure captions, scientific names of plants are not followed by authorship.

Examples of figure captions:

Figure 1Amaioua longipedicellata. A. Branchlet with male inflorescence. B. Branchlet with fasciculate female inflorescence. C. Fasciculate infructescence with immature fruits. D. trichotomous infructescence with mature fruits. E. Male flower in anthesis. F. Dissected male flower in anthesis. G. Female flower in anthesis. H. Dissected female flower in anthesis. A, C from Jardim & Delprete 6801 (CEPEC). B, E, F from Jardim & Delprete 6802 (CEPEC). D from photograph by J. Jardim (see fig. 2F). G, H from Jardim 5447 (CEPEC). Illustration by Piero Delprete.

Figure 2 – Morphological and anatomical characters of Synarthonia inconspicua. A–B. Thallus with ascomata. C. Section through ascoma. D. ascus. E. ascospores. A–E from Van den Broeck 6325 (BR). Scale bars: A = 500 μm; B = 250 μm; C = 50 μm; D–E = 10 μm.

Figure 4 – Gradation of differential edaphic variables (mean values) for the four ecotypes (G1, G2, G3 and G4) of Limonium spp. A. Sulphates (SO42- meq/l). B. Sand (%). C. Carbonates (CO32- %).

Alternatively, the capital letters are in bold between parentheses:

Figure 1 – Relationship between basal stem diameter (A) or flowering time (B) and number of seed pods for 2011 (red), 2012 (green), and 2013 (blue). Regression lines represent total linear selection.

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The pdf should contain all figures and their captions, all tables and their captions.

One figure per page, caption below.

Each table must start on a new page of the pdf.



Each table is a separate file, and is numbered according to its first citation in the text.

Each table has a complete caption at the top of the first page including the explanations of all abbreviations used in the table. The caption should be divided in two parts: (1) “Table + table number” followed by n-dash and a concise descriptive headline (in bold); (2) an explanatory part (in regular font), as in the following example:

Table 3Summarized results of two-factor ANOVA testing the effect of dune climate (Mediterranean-oceanic, M-subtropical and M-subdesert) and dune sector (fixed dunes vs. mobile and semi-mobile dunes) on soil nutrient content.
For each analysis, F-values are shown. LS, leaf size; SLA, specific leaf area; C, leaf [C]; N, leaf [N]; Ca, leaf [Ca]; K, leaf [K]; Mg, leaf [Mg];  P, leaf [P]; Fe, leaf [Fe]; Mn, leaf [Mn]; n.s. (not significant) P > 0.10; · P ≤ 0.10; * P ≤ 0.05; ** P ≤ 0. 01; *** P ≤ 0.001.

Text in table cells is typed using Times New Roman font, 9 pt. Capitals are avoided within table cells (exceptions: first letter of the heading of a row or a column, taxa names requiring a capital letter, such as family, genus, and some abbreviations, e.g. pH).

Please note that our journal does not use landscape format for the publication of papers. Tables in landscape format will be rotated 90 degrees counter-clockwise to fit a portrait page.

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Each figure is a separate file, and is numbered according to its first citation in the text.

All visual data (line diagrams, drawings, maps, graphs, photographs, plates...) are treated as figures.

Figure captions are provided separately (see Text File section, Figure captions).

Technical requirements

Figures are provided as high-quality files.

When possible (some graphs, simple line drawings), supply vector-based files. These files maintain high print-quality resolution at any size. Vector files are particularly appropriate for some data, such as phylogenetic trees. We can work with the following extensions: .svg, .ai, .pdf.

For raster images (photographs, complex line drawings, some maps...), supply the files at the appropriate dimensions to avoid image distortion. We can work with the following extensions: .tif, .jpg, .psd, .png

Figure size fits either single column size (85 mm wide) or double column size (up to 177 mm wide). Figure height is maximum 235 mm (preferably shorter to leave some space for the figure caption).

Line drawings and graphs are in black and white colour mode (bitmap) with a resolution of 1200 dpi.
Greyscale figures (e.g. photograph, map) are in greyscale colour mode with a resolution of minimum 450 dpi.
Colour images (RGB colour mode) are used where they enhance significantly clarity of the scientific information. They are of the same quality as the greyscale figures.

Electron and light micrographs have scale bars.

Figure formats

Formats accepted: .svg, .ai, .pdf (saved in high-quality), .tif, .jpg, .psd, .png

We cannot accept files in other formats; in particular, we cannot accept:
- MS PowerPoint files
- figures embedded in MS Word files
- figures prepared in PowerPoint or Word formats that have been converted to other, acceptable formats.

Figure composition and lettering

Related graphics are assembled into a single figure (composition) whenever possible. Each part (subfigure) of the composed figure is labelled A, B, C… and not treated as a separate figure. Labels are preferably placed in the upper left or lower left corner of the subfigure. Letters are in Arial or Calibri font, at the appropriate font size (depending on the figure height and width). White parting lines (2 mm wide) can be used to separate subfigures.

Titles or labels not absolutely necessary for understanding the figure should be removed and explained in the caption.



Additional data (not necessary to the basic understanding of the paper) or supporting data are supplied as separate supplementary file(s).

The number of supplementary files is limited to six per paper. Currently Plant Ecology and Evolution provides only DOI for papers, but not for supplementary files. Authors can decide to publish their supplementary files on a dedicated platform, e.g. Zenodo, and mention in their manuscript the Zenodo DOI links to access these additional files.

For a text file, a pdf with images, an Excel spreadsheet: the caption of a supplementary file must include the title of the manuscript and name(s) of its author(s), e.g. “Supplementary file 1 related to “title of the manuscript” (by names of authors)”.

For an image, a software file (e.g. nexus), a database file (e.g. csv), embed the title of the manuscript (and if possible authors names) in the metadata of the file (“properties” or “file info”).

Authors are free to group several items in one single supplementary file (e.g. several maps of a given area), but should provide a clear caption. The first part of the caption should be a general and short description for all items present in the supplementary file. The second part of the caption describes each item, and should be formatted like captions for composed figures (using letters A, B, C…).

The layout of the supplementary files is publication-ready.


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General requirements

Scientific names and their authorships follow the requirements specified in the Manuscript Text file instructions above.

Index Herbariorum acronyms are used for designation of herbaria.

Nomenclatural types are deposited in a public herbarium.

All references cited in the taxonomic treatment (valid publication of a basionym, of a synonym…) are included in the Reference section of the manuscript.

A list of taxonomic novelties present in the manuscript is provided at the end of the Text file.

Requirements for all taxonomic novelties (isolated/in frame of revision)

The following elements are required for any taxonomic novelty:
- an illustration (figure) clearly showing the diagnostic characters
- a comparison with related (or sympatric, or similar) taxa in a dichotomous key or table
- a discussion of the morphological characteristics, ecology, geography, or reproductive biology… that are the basis for its distinctiveness
- for names of new taxa, new combinations, names at new ranks, or replacement names designating organisms treated as fungi (including fossil fungi and lichen-forming fungi), authors are required to cite in the protologue the identifier issued by a recognized repository for the name (e.g. MycoBank, Index Fungorum, Fungal names).

Specific requirements for isolated taxonomic novelties

The journal accepts to publish isolated taxonomic novelties, e.g. a new species described outside the framework of a wider revision or monograph. Papers should be written in a way that they are attractive and interesting for a wider international audience. Such papers are mostly considered as short communications.

Manuscripts of this kind should include:
- a good introduction setting out the context of the higher taxon to which the novelty belongs, and making reference to recent taxonomic literature on the group – other taxonomic novelties recently published, etc.
- a sound discussion of the arguments for recognition of the new taxon; illustrations of the novelty
- some discussion about the area where the new species was found (climate, soil type, levels of endemism, status of collecting efforts in the region, other new species recently described from the area, conservation status of the area…)
- preferably, a conservation assessment (IUCN), or at least general statements on conservation.

For an example, the following paper can be downloaded: Darbyshire I. (2011) Barleria superata (Acanthaceae): a new suffruticose herb endemic to Burundi. Plant Ecology and Evolution 144: 367–371.     


Following the recommendation 32A of the International Code of Nomenclature for algae, fungi and plants, nomenclatural novelties should be introduced with the appropriate rank abbreviation, followed by the abbreviation ‘nov.’ (for novus, -a, -um).

Authors are required to use the following abbreviations in their treatment:
- family = fam.
- genus = gen.
- species = sp. (plural spp.)
- subspecies = subsp.
- variety = var.
- forma = f.
- synonym = syn.
- illegitimate name = nom. illegit.
- invalid name = nom. invalid.
- new combination = comb. nov.
- name at new rank = stat. nov.
- nomen nudum = nom. nud.


For emphasis, some terms are in bold in the taxonomic treatment:
- Introduction of a novelty: sp. nov., gen. nov., var. nov., subsp. nov., etc.
- Introduction of a new synonym: syn. nov.
- Introduction of a new combination, of a name at new rank: comb. nov., stat. nov.
- Introduction of illegitimate or invalid name: nom. illegit., nom. invalid.
- Designation of a lectotype, neotype, or epitype in the manuscript: designated here.

Treatment structure: novelties

Descriptions of new taxa include the following elements in the following order. Elements in italic are optional, depending on the information available. Third level headings are used to introduced all elements from 3 to 14 (e.g. “Type – ...”).

In accordance with article 38.1 of the International Code of Nomenclature for algae, fungi and plants, authors must provide either a diagnosis or a description for valid publication of the name, and are encouraged to provide both. 

1°) Name of the novelty (in bold italic), followed by the authorship, a comma, then the appropriate rank (in bold) and the abbreviation ‘nov.’ (in bold); e.g.
Synsepalum ntimii W.D.Hawthorne, sp. nov.

2°) Citation of figures and tables related to this taxon, e.g.
Figs 1, 2A, 3I–J, 4, tables 1, 2

3°) Diagnosis in Latin or English. The diagnosis is mandatory only if a full description is not provided. References to the illustrations can be added in the diagnosis.

4°) Type material citation. See the instructions for specimen citations.

5°) Full description of the new taxon, with major organs underlined. References to the illustrations can be added in the description. A diagnosis must be present if authors choose not to provide a full description.

6°) Distribution
7°) Chorology
8°) Habitat and ecology
9°) Phenology
10°) Vernacular names
11°) Uses
12°) Etymology
13°) IUCN conservation assessment. Recommended in the case of isolated novelty.
14°) Additional material examined
15°) Taxonomical notes or remarks.

Treatment structure: synonymy

For nomenclatural matters, homotypic synonyms are kept in one paragraph, followed by the type citation, and each heterotypic synonym is mentioned in a separate paragraph, followed by its own type citation. Heterotypic synonyms are listed in chronological order of publication. N-dashes (–) are used as separators within a paragraph.


Leplaea mayombensis (Pellegr.) Staner

Guarea mayombensis Pellegr. (Pellegrin 1921: 449, 1924: 54, 1939: 152); Pennington & Styles (1975: 495); Styles & White (1991: 43). – Leplaea mayombensis (Pellegr.) Staner (Staner 1941: 204); Staner & Gilbert (1958: 212). Type – Gabon, Tchibanga, 18 Jan. 1915, Le Testu 1990 (holotype: P; isotypes: BM, BR, E, K, WAG).
Leplaea coalescens Vermoesen (Vermoesen 1921: B64); Harms (1940: 137). – Guarea mayombensis Pellegr. var. coalescens (Vermoesen) Pellegr. (Pellegrin 1939: 152). – Leplaea mayombensis (Pellegr.) Staner var. coalescens (Vermoesen) Pellegr. (Staner 1941: 204). Type – D.R. Congo, Mayumba, Temvo, 26 Feb. 1919, Vermoesen 1680 (lectotype: BR, designated here).

New synonyms are indicated as syn. nov. at the end of the citation for the synonym.

Illegitimate and invalid names are indicated as nom. illegit. and nom. invalid. at the end of the citation for the name.

Treatment structure: new combination

New combination includes the basionym with a full and direct reference to its authorship and valid publication:

Leplaea cedrata (A.Chev.) E.J.M.Koenen & Wilde, comb. nov.

Trichilia cedrata A.Chev., Etudes Scientifiques et Agronomiques, Fascicule V: 214. 1909 (Chevalier 1909). – Guarea cedrata (A.Chev.) Pellegr. (Pellegrin 1928: 480); Pellegrin (1939: 154); Harms (1940: 135); Staner (1941: 187); Keay (Mar., 1958: 706); Staner & Gilbert (1958: 204); Aubréville (1959: 160); Voorhoeve (1965: 264); Pennington & Styles (1975: 495); Styles & White (1991: 43); Poorter et al. (2004: 418); Hawthorne & Jongkind (2006: 738). Type – Ivory Coast, Bouroukrou, s.d., A. Chevalier 16125 (lectotype: P; isolectotype: G, WAG). See also note 1.
Khaya canaliculata De Wild. (De Wildeman 1914: 377); De Wildeman (1920: 168). Type – D.R.Congo, Ganda-Sundi, 1913, J. de Briey 220 (holotype: BR).
Guarea alatipetiolata De Wild. (De Wildeman 1930: 71). Type – D.R.Congo, Eala, Nov. 1923, V.G. Goossens 4514 (holotype: BR).
Guarea kennedyi Burtt Davy, manuscript name; J.D. Kennedy (1930: 223), in obs., nom. nud. (see note 4).


When the lectotype (or neotype, or epitype) is designated in the manuscript itself:
(lectotype: BR, designated here).

When the lectotype (or neotype, or epitype) was designated before, the reference is needed:
(lectotype: AD, designated by Prescott 1984: 319).

Specimen citations

Specimen citations are grouped per country (one paragraph per country). Each citation is separated by a semicolon; each element in the citation is separated by a comma.

Authors are required to check carefully their specimen citations for consistency; for example, use always the same name for the same collector (except if different names designate two different collections with separate numbering system).

For 2 or more specimens sharing part of the data, use the expression “same data as preceding” and complete the citation with the differing data.

The elements are presented (when available) in the following order:
- country (before first citation)
- geopolitical unit(s) within country (e.g. state, region, district…), from largest to smallest
- locality where the specimen was collected, as written on the specimen’s label; if no locality was mentioned on the label, use the abbreviation ‘s. loc.’ (sine loco)
- geographical coordinates (see Style consistency in the Manuscript Text File section); authors are free to distinguish coordinates written on label and coordinates calculated by the authors; in this latter case, calculated coordinates should be between square brackets
- elevation as written on specimen’s label, e.g. “500 m”, “680 m a.s.l.”, “1400 ft”; if the elevation is not in metres, authors are free to add a conversion between square brackets
- collection date (see Style consistency in the Manuscript Text File section)
- phenology (fruit, flower…)
- collector(s) name (italic)
- collection number (italic); if no collection number, use “s.n.” without italic
- herbaria acronyms where specimen and duplicates are deposited; if the specimen was seen, do not add ‘!’; if the specimen was not seen, add ‘n.v.’; if the specimen was only seen as digital image, add ‘web’; authors are free to add the institutional barcode number, and in some cases, the url where the digital image is available.

Abbreviations in specimen citations

sine loco = s. loc.
sine dato = s.d.
sine collector = s. col.
sine numero = s.n.
fruit(s) = fr
open flower(s) = fl
flower bud(s) = fb
sterile = sterile
specimen not seen (non vidi) = n.v.
specimen only seen as digital image = web

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Privacy Statement

Privacy statement Plant Ecology and Evolution

  1. WHO ARE WE?

1.1 The controller

The following information is communicated to you so that you know the commitments regarding the protection of personal data made by Plant Ecology and Evolution, an academic journal co-owned by Agentschap Plantentuin Meise and by the Royal Botanical Society of Belgium (both organisations based in Belgium and governed by the Belgian law). Agentschap Plantentuin Meise acts as the controller for the processing of personal data referred to in this document.

1.2 Our Data Protection Officer

Agentschap Plantentuin Meise has designated a Data Protection Officer whose contact details are as follows:



In the context of processing of personal data, Agentschap Plantentuin Meise collects and processes the following data:

-        For authors: name, surname, institutional address, country, email address, IDs;

-        For reviewers: name, surname, institutional address, country, email address, IDs.



3.1 The purposes of our data processing

We are processing data for the following purposes:

-        The publication of the academic articles;

-        The creation of a database for its authors, reviewers and visitors of the website;

-        The implementation of the article review process;

-        The production of statistics;

-        The transmission of the metadata to libraries;

-        The transmission of data to online webpages and online indexing databases (IngentaConnect, Web of Science, Google Scholar, …)

3.2 The legal bases of our data processing

We implement data processing only if at least one of the following conditions is met:

-        your consent to the processing operations has been obtained;

-        the existence of our legitimate interest, or that of a third party, which justifies that we implement this processing of personal data;

-        the execution of a contract binding us to you requires that we implement this processing of personal data;

-        we are subject to statutory and regulatory obligations which require this processing of personal data.

3.3 The legitimate interests pursued

The legitimate interests pursued by Agenstschap Plantentuin Meise may consist in particular of:

-        Publication and dissemination of academic articles in Open Access

-        Peer-review of academic articles



The personal data we collect, and those we obtained subsequently, are intended for us in our capacity as controller.

We ensure that only authorised persons have access to this data. Our service providers can be recipients of this data to perform the services we entrust to them. Some personal data may be sent to third parties or to legally authorised authorities in order to meet our legal, regulatory or contractual obligations.

They may be communicated to these entities for the purposes referred to in this privacy policy. These operations are carried out on the basis of instruments that comply with applicable regulations and are capable of ensuring that your rights are protected and respected.



We process and store your personal data in Belgium. We transfer your personal data to partners in the following countries:

-        Canada.

-        United States.

Each of these transfers is governed by legal instruments that comply with the applicable legal framework. The following countries [Canada and United States] benefit from an adequacy decision, which means that they offer your personal data a level of protection equivalent to the one which is applied on the European Union territory. Transfers made to other countries are covered by the following appropriate safeguards:

-        standard contractual clauses approved by the European Commission.



The periods for which we keep your personal data are proportionate to the purposes for which your data were collected. Our data storage policy is organized as follows:

-        For data concerning authors: as long as the article is published;

-        For data concerning reviewers: as long as necessary: due to potential scientific fraud cases, the data might be saved for 25 years after the publication or rejection of a paper.



7.1 Your right of information

You acknowledge that this privacy policy provides you with information about the purposes, legal framework, interests, recipients or categories of recipients with whom your personal data are shared, and the possibility of a data transfer to a third country or international organisation.

In addition to this information and with the aim of ensuring fair and transparent processing of your data, you further acknowledge that you have received additional information concerning the period for which your personal data will be kept.

If we decide to process data for purposes other than those indicated, all information relating to those new purposes will be communicated to you.

7.2 Your right of access to and rectification of your data

You have the right to access and rectify your personal data, which you can exercise with our Data Protection Officer at the following email address:

In this respect, you have the confirmation as to whether or not your personal data are being processed and where this is the case, access to your data and the following information:

-        the purposes of the processing;

-        the categories of personal data concerned;

-        the recipients or categories of recipient as well as the international organisations to whom the personal data have been or will be disclosed, in particular recipients in third countries;

-        where possible, the envisaged period for which the personal data will be stored, or, if not possible, the criteria used to determine that period;

-        the existence of the right to request from the controller rectification or erasure of personal data or restriction of processing of personal data concerning the data subject or to object to such processing;

-        the right to lodge a complaint with a supervisory authority;

-        where the personal data are not collected from the data subjects, any available information as to their source;

-        the existence of automated decision-making, including profiling, and in this case, meaningful information about the logic involved, as well as the significance and the envisaged consequences of such processing for the data subject.

You can ask us to, as the case may be, rectify or complete your personal data that are inaccurate, incomplete, equivocal or expired.

7.3 Your right to erasure of your data

You can ask us to erase your personal data where one of the following grounds applies:

-        the personal data are no longer necessary in relation to the purposes for which they were collected or otherwise processed;

-        you withdraw the consent you have previously given;

-        you object to the processing of your personal data and there is no legal reason for such processing;

-        the processing of personal data does not comply with the provisions of the applicable legislation and regulations;

-        your personal data have been collected in relation to the offer of information society services to children under 16 years of age.

Nevertheless, the exercise of this right will not be possible when the retention of your personal data is necessary for compliance with statutory or regulatory provisions and in particular for example for the establishment, exercise or defence of legal claims.

7.4 Your right to restriction of processing

You may request restriction of processing of your personal data in the cases provided for by law and regulation.

7.5 Your right to object to data processing

You have the right to object to the processing of personal data concerning you when the processing is based on the legitimate interest of the controller.

7.6 Your right to data portability

From 25 May 2018, you have the right to portability of your personal data.

The data on which this right can be exercised are:

-        only your personal data, which excludes anonymized personal data or data that does not concern you;

-        declarative personal data and personal data relating to our functioning, as mentioned above;

-        personal data which do not adversely affect the rights and freedoms of others such as those protected by trade secrets.

This right is limited to processing based on consent or contract as well as to personal data that you have personally generated.

This right does not include derived or inferred data, which are personal data created by Agenstschap Plantentuin Meise.

7.7 Your right to withdraw your consent

When the data processing we carry out is based on your consent, you may withdraw it at any time. We will then stop processing your personal data but this will have no impact on the previous transactions to which you have consented.

7.8 Your right to lodge a complaint with a supervisory authority

You have the right to lodge a complaint with the Belgian Data Protection Authority ( on the Belgian territory without prejudice to any administrative or judicial remedy.

7.9 Your right to define post-mortem directives

You can give instructions in relation to the storage, erasure and communication of your personal data after your death to a certified trusted third party in charge of enforcing the wishes of the deceased in compliance with the applicable legal framework.           

7.10 How to exercise your rights?

All the rights enumerated above can be exercised by sending a request, together with a copy of an ID document, to the Data Protection Officer (

Regarding the exercise of your right of information, we are not required to provide you with information where:

-        you already possess the information;

-        the recording or disclosure of your personal data is expressly laid down by law;

-        the provision of information to you proves to be impossible;

-        the provision of information to you would involve a disproportionate effort.